The article is interesting and there is some debate, although it would seem to lean toward "yes."
The most interesting part is the guy who logically states, "It is a huge mistake to think studying homosexuality in animals gives us a greater understanding of human behavior." Although that is a bit presumptuous.
What he really should have said is that studying animal socializing does not tell us about human social behavior. Although it can lead to a better understanding of how biology effects behavior. It's important to understand that there is a distinction and that it is often blurred.
Studying the biological imperatives of homosexuality doesn't dictate how it will be defined or viewed or otherwise relate to human society, because human society is the product of imagination, ethics, morality, logic and other such mental constructs of individuals that have little to do with species biology.